Evolutionary Tree of Arthropods

Note that Mya is million years ago. For example, "48.6～40.4 Mya" shows a range between 48.6 and 40.4 million years ago. The time range and the epoch are shown in the Geological Time Table.

Phylogenetic Tree of Arthropods

Kingdom Animalia
- Phylum Arthropod
  - Class Dinocaridida (Anomalocaris, Opabinia; 530～505 Mya)
    - Order Radiodonta
  - Class Megacheira (Crustacean-like arthropods; 542～501 Mya)
  - Subphylum Trilobitomorpha (Trilobites, Marrella)
    - Class Marrellomorpha
      - Order Marrelliina
    - Class Trilobita (Trilobites; 542～251 Mya)
      - Order Agnostida (530～383.7 Mya) Trilobite
      - Order Nectaspida Trilobite
      - Order Redlichiida (530～488.3 Mya) Trilobite
      - Order Corynexochida (520～376.1 Mya) Trilobite
      - Order Lichida (503～376.1 Mya) Trilobite
      - Order Phacopida (488.3～345.3 Mya) Trilobite
      - Subclass Librostoma
        
        Order Proetida Trilobite
        Order Asaphida (516～418.1 Mya) Trilobite
        Order Harpetida (513～376.1 Mya) Trilobite
        Order Ptychopariida (530～422.9 Mya) Trilobite
  - Subphylum Chelicerata (Horseshoe crabs, Scorpions, Spiders, Mites)
    - Class Arachnida (Spiders, Scorpions)
      - Order Trigonotarbida (418.7～299 Mya)
      - Order Amblypygi (Whip spiders)
      - Order Araneae (Spiders; 409.1～0 Mya)
        
        Suborder Mesothelae (Basal spiders) Spider
        Suborder Opisthothelae (Spiders)
        
        Infraorder Araneomorphae (Most common spiders) Spider
        Infraorder Mygalomorphae (Tarantulas, Tarantula-like spiders) Spider
      - Order Opiliones (Phalangids, Harvestmen; 336 Mya～)
      - Order Palpigradi (Microwhip scorpions)
      - Order Pseudoscorpionida (False scorpions)
      - Order Ricinulei (Hooded tickspiders)
      - Order Schizomida (Shorttailed whipscorpions)
      - Order Scorpiones (Scorpions) Scorpion
      - Order Solifugae (Sun spiders)
      - Order Thelyphonida (Vinegarroons)
    - Class Acari (Mites, Ticks)
      - Order Holothyrida
      - Order Ixodida (Ticks)
      - Order Mesostigmata (Bird mites, Phytoseiid mites)
      - Order Trombidiformes (Plant parasitic mites)
      - Order Sarcoptiformes
      - Order Oribatida (Oribatid mites)
    - Class Merostomoidea (Intermediate form between trilobites and chelicerates)
      - Order Limulavida
    - Class Merostomata (Horseshoe crabs, Sea scorpions)
      - Order Eurypterida (Sea scorpions; 460.5～258 Mya) Sea scorpion
      - Order Xiphosura (Horseshoe crabs; 542 Mya～) Horseshoe crab
      - Order Aglaspida (542～488.3 Mya)
    - Class Pycnogonida
      - Order Pantopoda (Sea spiders)
  - Subphylum Myriapoda (Millipedes, Centipedes)
    - Class Arthropleuridea
      - Order Arthropleurida
    - Superclass Opisthogoneata
      - Class Chilopoda (Centipedes)
        
        Order Scutigeromorpha (House centipedes)
        Order Lithobiomorpha (Stone centipedes)
        Order Craterostigmomorpha
        Order Scolopendromorpha (Tropical centipedes)
        Order Geophilomorpha (Soil centipedes)
    - Superclass Progoneata
      - Class Diplopoda (Millipedes; 318.1 Mya～)
        
        Order Polyxenida
        Order Glomerida (Pill millipedes)
        Order Sphaerotheriida (Giant pill millipedes)
        Order Polydesmida (Polydesmid millipedes)
        Order Chordeumatida
        Order Julida (Julid millipedes)
        Order Spirobolida
        Order Spirostreptida
      - Class Pauropoda
        
        Order Pauropodina (Pauropods)
      - Class Symphyla
  - Subphylum Hexapoda (Insects, others)
    - Class Insecta (Insects)
      - Subclass Apterygota (Primitive wingless insects)
        
        Order Monura
        Order Archaeognatha (Bristletails)
        Order Thysanura (Silverfish)
      - Subclass Pterygota (Winged insects)
        
        Infaclass Paleoptera (New winged insects)
        
        Order Ephemeroptera (Mayflies)
        Order Palaeodictyoptera (Six-winged insects; 318.1～216.5 Mya)
        Order Megasecoptera (307.1～268 Mya)
        Order Protodonata (Giant dragonflies; 318.1～199.6 Mya) Dragonfly
        Order Odonata (284 Mya～)
        
        Suborder Anisozygoptera Dragonfly
        Suborder Anisoptera (Dragonflies) Dragonfly
        Suborder Zygoptera (Damselflies) Dragonfly
        
        Infaclass Neoptera (Old winged insects)
        
        Superorder Exopterygota
        
        Order Protorthoptera (318.1～216.5 Mya)
        Order Plecoptera (Stoneflies; 268 Mya～)
        Order Embioptera (Webspinners)
        Order Zoraptera (Angel insects)
        Order Dermaptera (Earwigs; 167.7 Mya～)
        Order Orthoptera (Crickets, Grasshoppers; 307.1 Mya～)
        
        Superorder Dictyoptera
        
        Order Phasmatodea (Stick insects, Leaf insects; 183.7 Mya～)
        Order Notoptera (Ice-crawlers, Gladiators)
        Order Blattaria (Cockroaches; 307.1 Mya～)
        Order Isoptera (Termites; 150.8 Mya～)
        Order Mantodea (Mantis) Mantis
        
        Superorder Paraneoptera
        
        Order Psocoptera (Booklice, Barklice)
        Order Thysanoptera (Thrips; 284 Mya～)
        Order Phthiraptera (Lice)
        Order Hemiptera (True bugs; 284 Mya～)
        
        Suborder Auchenorrhyncha
        
        Infraorder Cicadomorpha (Cicadas, Leafhoppers, Treehoppers)
        Infraorder Fulgoromorpha (Planthoppers)
        
        Suborder Coleorrhyncha (Moss bugs, Beetle bugs)
        Suborder Heteroptera (True bugs)
        
        Infraorder Cimicomorpha (Assassin bugs, Bed bugs, allies)
        Infraorder Dipsocoromorpha
        Infraorder Enicocephalomorpha (Unique-headed bugs)
        Infraorder Gerromorpha (Water treaders)
        Infraorder Leptopodomorpha (Shore bugs)
        Infraorder Nepomorpha (True water bugs)
        Infraorder Pentatomomorpha (Stinkbugs)
        
        Suborder Sternorrhyncha (Aphids, Whiteflies, Scales)
        
        Superorder Endopterygota
        
        Order Hymenoptera (Ants, Bees; 228 Mya～)
        
        Suborder Apocrita
        
        Infraorder Aculeata (Honeybees, Ants, Hornets)
        Infraorder Parasitica (Parasitoid wasps)
        
        Suborder Symphyta (Sawfly)
        
        Order Coleoptera (Beetles; 284 Mya～)
        
        Suborder Adephaga (Ground beetles, Diving beetles)
        Suborder Archostemata
        Suborder Myxophaga
        Suborder Polyphaga
        
        Infraorder Bostrichiformia (Furniture beetles, Skin beetles)
        Infraorder Cucujiformia (Longhorn beetles, Lady beetles, Weevils) Lady beetle
        Infraorder Elateriformia (Fireflies) Firefly
        Infraorder Scarabaeiformia (Stag beetles, Scarab beetles, Rhinoceros beetles) Rhinoceros beetle
        Infraorder Staphyliniformia
        
        Order Strepsiptera (Twisted-winged parasites; 112 Mya～)
        
        Superorder Neuropterida
        
        Order Raphidioptera (Snakeflies; 161.2 Mya～)
        Order Megaloptera (Alderflies)
        Order Neuroptera (Net-veined insects, Lacewing; 260.5 Mya～)
        
        Superorder Antliophora
        
        Order Mecoptera (Scorpionflies; 284 Mya～)
        Order Siphonaptera (Fleas; 112 Mya～)
        Order Diptera (True flies; 228 Mya～)
        
        Suborder Nematocera (Mosquitoes, Crane flies, Gnats, Midges)
        Suborder Brachycera　(House flies, Horse flies)
        
        Superorder Amphiesmenoptera
        
        Order Trichoptera (Caddisflies; 268 Mya～)
        Order Lepidoptera (Butterflies, Moths)
        
        Suborder Aglossata (Moths) Moth
        Suborder Glossata
        
        Infraorder Heteroneura (Moths, Swallowtail butterflies, Four-footed butterflies) Butterfly Moth
        Infraorder Dacnonypha (Moths) Moth
        Infraorder Lophocoronina (Moths) Moth
        Infraorder Exoporia (Moths) Moth
        Infraorder Neopseustina (Moths) Moth
        
        Suborder Heterobathmiina (Moths) Moth
        Suborder Zeugloptera (Moths) Moth
    - Class Entognatha
      - Order Collembola (Springtailes)
      - Order Diplura (Two-pronged bristletails)
      - Order Protura (Coneheads)
  - Subphylum Crustacea (Crabs, Lobsters, Crayfish, Shrimp, Krill, Barnacles)
    - Class Thylacocephala (513～161.2 Mya)
      - Order Concavicarida
    - Class Branchiopoda (Brine shrimp; 520 Mya～)
      - Order Anostraca (Fairy shrimps)
      - Order Notostraca
      - Order Laevicaudata (Clam shrimps)
      - Order Cladocera (Water fleas)
    - Class Remipedia
      - Order Nectiopoda
    - Class Cephalocarida (Horseshoe shrimp)
      - Order Brachypoda
    - Class Maxillopoda (Barnacles, Fish lice)
      - Subclass Thecostraca
        
        Order Kentrogonida
        Order Akentrogonida
        Order Pedunculata (Goose barnacles)
        Order Sessilia (Acorn barnacles)
      - Subclass Branchiura (Fish lice)
        
        Order Arguloida
      - Subclass Pentastomida (Tongue worms)
        
        Order Cephalobaenida
        Order Porocephalida
      - Subclass Mystacocarida
        
        Order Mystacocaridida
      - Subclass Copepoda
        
        Order Calanoida
        Order Cyclopoida
        Order Gelyelloida
        Order Harpacticoida
        Order Monstrilloida
        Order Poecilostomatoida
        Order Siphonostomatoida
    - Class Ostracoda (Seed shrimps; 530 Mya～)
      - Subclass Myodocopa
        
        Order Myodocopida
        Order Halocyprida
      - Subclass Podocopa
        
        Order Platycopida
        Order Podocopida
    - Class Malacostraca (Lobsters, Crabs, Shrimps; 516 Mya～)
      - Subclass Phyllocarida
        
        Order Canadaspidida (516～501 Mya)
        Order Leptostraca
      - Subclass Hoplocarida
        
        Order Stomatopoda (Mantis shrimp; 339.4 Mya～)
      - Subclass Eumalacostraca
        
        Superorder Syncarida
        
        Order Palaeocaridacea
        Order Bathynellacea
        Order Anaspidacea
        
        Superorder Peracarida
        
        Order Spelaeogriphacea
        Order Thermosbaenacea
        Order Lophogastrida
        Order Mysida (Opossum shrimp)
        Order Mictacea
        Order Amphipoda
        Order Isopoda (Woodlice, Slaters; 272.5 Mya～)
        Order Tanaidacea
        Order Cumacea (Hooded shrimp)
        
        Superorder Eucarida
        
        Order Euphausiacea (Krill)
        Order Amphionidacea
        Order Decapoda (Crabs, Lobsters, Shrimp; 376.1 Mya～)
        
        Suborder Dendrobranchiata (Prawns) Shrimp
        Suborder Pleocyemata
        
        Infraorder Polychelida Shrimp
        Infraorder Glypheidea Shrimp
        Infraorder Stenopodidea Shrimp
        Infraorder Caridea (Shrimps) Shrimp
        Infraorder Achelata (Lobsters) Shrimp
        Infraorder Astacidea (Lobsters) Shrimp
        Infraorder Thalassinidea (Mud lobsters) Shrimp
        Infraorder Anomura (Sand crabs) Crab
        Infraorder Brachyura (Crabs) Crab